On the systematics of the Anomalina subtribe in Southeast Asia (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)
DOI:
https://doi.org/10.33910/2686-9519-2021-13-4-581-594Keywords:
scarab beetles, taxonomic descriptions, new synonymies, identification keys, China, Indo-China, MyanmarAbstract
Anomala hamuliphalla (Myanmar: Kachin State, Laos), A. macrorhynchos (China: Yunnan) and A. malivirens (Vietnam) spp. nov. are described. Anomala graminea Ohaus, 1905 is allocated into the genus Tonkinilla gen. nov. The following new synonyms are established: Choumala Kobayashi, 2008 = Anomala Samouelle, 1819; Choumala choui Kobayashi, 2008 = Anomala phalaena Ohaus, 1915. Anomala perplexa diana Zhang et Lin, 2008 is raised to a full species. The limits of the species group of Anomala dalmanni are discussed and a key for identification is given.
Diagnoses of new taxa. Anomala hamuliphalla sp. nov. is unique in the shape of the left paramere extended into a hook, and in the bifurcate distal tip of the basal plate of the aedeagus with the rami divergent.
Anomala macrorhynchos sp. nov. is similar to A. millestriga and A. punctatissima, but can be distinguished in the shape of the parameres.
Anomala malivirens sp. nov. is similar to A. diana, but differs in the shorter parameres with more rounded apices in profile.
Tonkinilla gen. nov. (type species: Anomala graminea Ohaus, 1905) is similar to Mimela in the presence of the prosternal process, but the latter is compressed anteroposteriorly instead of laterally. The new genus can be further distinguished from the known genera by the peculiar pilosity, exposed labrum, loss of the elytral epipleura and the shape of the aedeagus.
References
Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Pt II (Rutelinae, Desmonycinae, and Euchirinae). London: Taylor & Francis Publ., 387 p. (In English)
Blanchard, E. (1851) Catalogue des collections Entomologiques du Muséum d’Histoire naturelle de Paris. Classe des Insectes, Ordre des Coléoptères I. Paris: Gide et Baudry Publ., pp. 129–240. (In French)
Kim, J.-I. (1998) Taxonomic study of Korean Rutelidae VI. Two new genera and some removal species from Korean Anomala. The Korean Journal of Entomology, vol. 28, no. 4, pp. 311–316. (In English)
Kobayashi, H, Chou, W.-I. (2008) Description of a new genus of Anomalini and two new species of Hoplini and Melolonthini from Taiwan, with two new records of scarabaeid beetles. Kogane, vol. 9, pp. 69–76. (In English)
Kobayashi, H., Fujioka, M. (2013). Notes on the genus Pseudosinghala Heller from Southeast Asia. Kogane, vol. 14, pp. 85–92. (In English)
Krajcik, M. (2012). Checklist of the world Scarabaeoidea. Animmax. X, Supplement 5, pp. 1–278. (In English)
Machatschke, J. W. (1957) Coleoptera Lamellicornia Fam. Scarabaeidae Subfam. Rutelinae. Tribus. Anomalini. Zweiter Teil. In: P. A. G. Wytsman (ed.). Genera insectorum. Fascicule 199 (B). Brussels: Desmet-Verteneuil Verlag, pp. 1–219. (In German)
Ohaus, F. (1905) Beiträge zur Kenntnis der Ruteliden. Deutsche Entomologische Zeitschrift, vol. 1, pp. 81–99. (In German)
Ohaus, F. (1915) XVII Beitrag zur Kenntnis der Ruteliden. Stettiner entomologische Zeitung, vol. 76, pp. 88–143. (In German)
Ohaus, F. (1943) Revision der Gattung Mimela Kirby. Deutsche entomologische Zeitschrift, vol. 75, pp. 65–88. (In German)
Paulian, R. (1958) Coléoptères Scarabéides de L’Indochine (Rutélines et Cétonines). Annales de la Société entomologique de France, vol. 127, pp. 73–105. (In French)
Paulian, R. (1959) Coléoptères Scarabéides de L’Indochine (Rutélines et Cétonines) (Suite). Annales de la Société entomologique de France, vol. 128, pp. 35–136. (In French)
Prokofiev, A. M. (2012) Dva novykh vida roda Anomala Sam. iz tsentral’nogo V’etnama (Coleoptera: Scarabaeidae) [Two new species of the genus Anomala Sam. from Central Vietnam (Coleoptera: Scarabaeidae)]. Russian Entomological Journal, vol. 21, no. 4, pp. 385–393. (In Russian)
Prokofiev, A. M. (2013) A new genus of Anomalini from Vietnam (Coleoptera: Scarabaeidae: Rutelinae). Russian Entomological Journal, vol. 22, no. 1, pp. 5–7. (In English)
Prokofiev, A. M. (2015) New Anomala Samouelle, 1819 from South-East Asia (Coleoptera: Scarabaeidae: Rutelinae). Russian Entomological Journal, vol. 24, no. 1, pp. 37–59. https://doi.org/10.15298/rusentj.24.1.04 (In English)
Ratcliffe, B. C., Jameson, M. L., Zorn, C. (2018) Ganganomala saltini Ratcliffe, Jameson, and Zorn, a new genus and species of Anomalini (Coleoptera: Scarabaeidae: Rutelinae) from Bangladesh and Nepal, with a revised circumscription of the tribe. The Coleopterists Bulletin, vol. 72, no. 4, pp. 717–735. https://doi.org/10.1649/0010-065X-72.4.717 (In English)
Zhang, B., Lin, P. (2008) The Anomala sinica species group from China. Oriental Insects, vol. 42, no. 1, pp. 125–141. https://doi.org/10.1080/00305316.2008.10417539 (In English)
Zorn, C., Kobayashi, H., Wada, K. (2017) Notes on the genus Anomala Samouelle, 1819 in Vietnam and neighboring regions: Eight new species and faunistic records. Beiträge zur Entomologie, vol. 67, no. 2, pp. 325–352. https://doi.org/10.21248/contrib.entomol.67.2.325-352 (In English)
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